resilin
a protein that fleas use to jump
what do humans use resilin for?
artificial heart valves, spinal discs, tennis shoes
why are arthropods so diverse and abundant?
versatile exoskeleton2)jointed appendages and striated muscles3) oxygen is piped directly into tissues and cells4) highly developed sensory organs5) complex behavioural patterns6) metamorphosis
Arthropod characteristics
Jointed limbs- Exoskeleton- Heart with Ostia- Trachea- Compound eyes- Malpighian tubules- Mandibles- No cilia at any life stage- Tagmatization: allows functional specialization of 'groups offused & modified segments'
exoskeleton layers
thin outer epicuticle and inner thicker procuticle
epicuticle
The outermost layer of the insect cuticle designed to prevent water loss. impermeable to water
procuticle
inner thick layer of an arthropod shell that consists primarily of layers of protein and chitin
chitin
tough and resilient but flexible
sclerotization
cross-linking of proteins which darkens the cuticle
ecdysis (molting)
Shedding of the cuticle and growing a new, larger cuticle after the body rapidly enlarges.
how many moults does it take before an animal becomes and adult?
4-7
Arthropods locomotion
because they are incased in a suit of rigid plates, the coelom fluid (haemocoel) plays NO major role in locomotion and is reduced- arthropods typically move by walking, swimming, or flying
arthropod circulatory system
open circulatory system- ostium
ostium
allows blood to enter heart from coelom but prevent it from leaving before it reaches the front
gills
outgrowths of the skin/modified appendages covered by thin, but permeable exoskeleton
book gill location
external
book lung location
internal, and connected by spiracles- maximizes gas exchange
tracheal system in a terrestrial arthropod
complex network of tubules that deliver oxygen directly to the animals tissues and cells
do arthropods carry their oxygen in haemolymp?
no, but vertabrates do .
spiracles
how the trachea is connected to the internal environment- can open and close to reduce water loss.
can velvet worms open and close their spiracles?
NO, they are always open.
do terrestrial arthropods transport oxygen in blood?
no. they lack haemolymph to bind oxygen
aquatic arthropod respiration
they use gills and transport oxygen in their blood, use hemocyanin
3 distinct regions in arthropod digestion
foregut, midgut, hind gut
foregut
highly muscular, breaks down food into small pieces to increase surface area
midgut
secretes enzymes to digest food and also absorbs nutrients
hind gut
reabsorbs water
arthropod excretion
solid wastes via anus
some aquatic arthropods excrete through
gills and digestive glands
some crustaceans have
green (antennal) glands- connected to pores under antennae
coxal glands
excretory organs of spiders
Malpighian tubules
found in most insects, arachnids, and myriapods
all arthropod muscles are:
striated
striated muscles
allow for voluntary muscle movements.- are attached to skeleton- have more mitochondria
Why are striated muscle cells so important for arthropods?
allows for much more rapid muscle contractions, this allows flight to be possible!
arthropod brain
supraesophageal ganglion
supraesophageal ganglion 3 regions
protocerebrum, deutocetebrum, tritocerebrum
protocerebrum
vision
deutocetebrum
processes sensory info from antennae
tritocerebrum
integrates sensory inputs from the protocerebrum and deutocerebrum
what's located immediately behind the arthropod brain?
subesophageal ganglion
subesophageal ganglion
part of the ventral nerve cord, it's connected to the brain
controls sensory organs and muscles of the mouthparts and salivary glands.
how are anterior ganglia connected?
by a pair of ventral ganglionate nerve cord running along the length of the abdomen
why are spiders more centralized?
they have no ganglia in abdomen or terminal
Arthropod sensory
intricate mouthparts, statocysts, and antennae sensitive hairs on body can detect touch, water currents, or chemicals
statocysts
sensory cells that help determine direction of gravity
arthropod vision
they use ocelli and compound eyes insects usually have 2 compound eyes and 3 ocelli
ocelli
non image forming, can only detect direction of light
how many ocelli do arachnids have?
typically 8. a few species have good eye sight.
ommatidia
compound eyes- hexagonal visual units 10x loner than wide.
the cornea and crystalline cone, together focus the light onto the..
rhabdom
light energy absorbed by rhabdom is converted via..
photo receptor cells, into nerve signals the brain can understand.
# of ommatidium
ranges from 15- several thousand
what animal has the most complex eyes?
the mantis shrimp.has at least 12 different photoreceptors.
earliest identifiable fossils of land animals...
arthropod fossils, 419ma
how did jointed exoskeleton and limbs make arthropods pre adapted for land?
provided protection against desiccation, support against gravity, and means of locomotion that wasn't dependant on water.
arthropod phylogeny
thought to be monophyletic
2 arthropod clades
chelicerate and mandibulata
Chelicerata key features
pointed, fang like appendages, lacks jaws; used to grasp food. considered a subphyla
mandibulata key feature
jaw like; used for chewing; divided into 3 extant subphyla
3 subphyla of mandibulata
myriapods, crustacea, hexapods
subphyla trilobita
extinct. no know descendants- had eyes of modern bees
examples of chelicerata
horseshoe crabs, spiders, ticks, mites, scorpions, sea spiders.
Chelicerata characteristics
suck liquid from prey- no antennae-segmented (lost in acari)- no distinct head, 2 distinct portions the segments are organized into (tagmata)- 6 pairs of cephalothoracic appendages
2 distinct portions in chelicerata
cephalothorax- fused head and thorax; called prosoma- abdomen (opisthosoma)
6 pairs of cephalothoracic appendages in chelicerata
1 pair of chelicerae (fang like appendages, adjacent mouth)1 pair of pedipalps (in front of head)4 pairs of walking legs.
what is the function of pedipalps in arachnids?
functions as an organ of touch and reproduction.
3 arthropod classes
merostomata (horse shoe crabs), pychnogonida (sea spiders), arachnida (spiders, scorpions, mites, ticks)
Merostomata Characteristics
long tail- used as rudder to help flip itself overbook gills- respiration. mouth between their legs- 10 eyes
2 subclasses of merostomata
eurypterida and xiphosurida
eurypterida
extinct form of giant water scorpions.- may be longest arthropod
xiphosurida
horseshoe crabs- not a crab or crustacean.- only 5 species left. behaviour based on mood and tidal cycles.- 10 eyes and can see uv light.
horseshoe crab feeding
small clams, crustaceans, worms.
crush hard food between their legs before passing it to their mouth.
horseshoe crab eggs
can lay 120 000 eggs a season. take 2 weeks to hatch.
how often do horseshoe crab larvae moult?
6 times during their first year. 18x before they reproduce.
when can horseshoe crabs reproduce.
at 9-11 years old
class pychnogonida
sea spiders- not true spiders or even arachnids.
class pychnogonida form
-mostly cephalothorax- abdomen reduced to a short stump.- variable number of walking legs
why is class pychnogonida proboscis unique?
it has chelicerae at terminal- allows them to suck juices from soft-bodied animals.
class pychnogonida speed
Very slow moving
class pychnogonida feeding
feed on immobile prey such as sponges and sea anemone
how many simple eyes in class pychnogonida ?
they have 2-4 simple eyes
in class pychnogonida, genitals and digestive system...
extend well into legs
class pychnogonida use their legs as gills and...
their guts ate hearts.
5 major orders of arachnids
araneae (spiders) 2)scorpionida (scorpions) 3)solpugida (sun or caramel spiders) 4)opiliones (harvestmen) 5)acari (ticks and mites)
in araneae, magmata are joined by a narrow...
pedicel
araneae feeding
ALL predaceous and largely feed on insects
araneae chelicerae
functions as fangs and bear ducts from venom glands.
araneae segmentation
MOST do no have external segmentation on abdomen
Liphistiidae
the araneae the does have segmentation
Araneae Excretion
up to 4 pairs of coxal glands and Malpighian tubules for excretion
araneae respiration
have book lungs, trachea, or both. ( other arachnid orders use only trachea)
spinnerets
organ in spiders that contains silk glands
silk is stronger than
steel of the same diameter
not all spiders spin webs...
but ALL produce silk, it has many other uses -acts as a balloon underwater for air.
red banded crab spider
awaits prey on flowers to which they are matched (they camouflage)
araneae reproduction
male spins web, deposits drop of sperm on it.- inserts a pedipalp into females genital opening- female fertilizes egg in silken cocoon which she might carry or attach to a plant
where is sperm stored in araneae
in his pedipalps
how many eggs in a araneae cocoon
hundres. they hatch in around 2 week
black widows are know for
cannibalizing their mates but this doesnt actually happen all the time